現生人類の拡散とは? わかりやすく解説

Weblio 辞書 > 辞書・百科事典 > 百科事典 > 現生人類の拡散の意味・解説 

現生人類の拡散

(Early human migrations から転送)

出典: フリー百科事典『ウィキペディア(Wikipedia)』 (2024/03/26 15:13 UTC 版)

現生人類の拡散(げんせいじんるいのかくさん)では、ホモ・サピエンスが世界各地へ拡散した経緯について述べる。ホモ・サピエンスが誕生し、移動を始めたのはおおよそ30万年前であるといわれている[note 1][3]アフリカ単一起源説では、おおよそ7万年前から5万年前に東アフリカを発ったホモ・サピエンスが現世人類の祖であるとされており、現生人類がアフリカを出てアジア南沿岸経由で近オセアニア地域への移動を始めたのがおおよそ7万年から5万年前とされており(人類の南方地域への分散英語版)、約4万年前までにヨーロッパ中に分散した。ただし、それ以前にもアフリカを出たホモ・サピエンスが存在する。イスラエルから19万4千年–17万7千年前のホモ・サピエンスの化石、ギリシャから約21万年前のホモ・サピエンスの化石が見つかっているが、いずれもそれ以前に定住していたネアンデルタール人との生存競争に敗れたと考えられている[4][5][6][7][3]


注釈

  1. ^ Schlebusch et al.によれば、南アフリカの古代人のゲノム解析によって現代人が26万年前から35万年前には誕生していたと推定されている[1][2] (archaic admixture).
  2. ^ 現生人類における旧人類の遺伝子の割合は、祖先がネアンデルタール人と混血したユーラシアの人々で推定1-4%(Lohse (2014) によれば3.4–7.3%[9])、Reich et al. (2010) によれば祖先がデニソワ人と混血したメラネシアの人々で推定4–6%[10]、Hammer et al. (2011) によれば未知の旧人類と混血したサブ・サハラの狩猟採集民族で推定2%とされる[11]
  3. ^ Estimated split times given in the source cited (in kya): Human-Neanderthal: 530–690, Deep Human [H. sapiens]: 250–360, NKSP-SKSP: 150–190, Out of Africa (OOA): 70–120.[1]
  4. ^ この変異は、汗腺や歯、髪の毛の太さ、胸部組織の形質に影響を及ぼす[92][93]。また、東アジア人の遺伝子には選択圧が示唆される対立遺伝子が多数みられる。これらの遺伝子は特にEDAR、ADH1B、ABCC1、ALDH2の遺伝子に関係する。その内、1万1千年前以降から現れた東アジア型のADH1B遺伝子は稲作に関連しているという[94]

出典

  1. ^ a b Schlebusch (3 November 2017). “Southern African ancient genomes estimate modern human divergence to 350,000 to 260,000 years ago”. Science 358 (6363): 652–655. Bibcode2017Sci...358..652S. doi:10.1126/science.aao6266. PMID 28971970. 
  2. ^ Stringer, C (2012). “What makes a modern human”. Nature 485 (7396): 33–35. Bibcode2012Natur.485...33S. doi:10.1038/485033a. PMID 22552077. 
  3. ^ a b c Harvati, Katerina (10 July 2019). “Apidima Cave fossils provide earliest evidence of Homo sapiens in Eurasia”. Nature 571 (7766): 500–504. doi:10.1038/s41586-019-1376-z. PMID 31292546. 
  4. ^ Callaway, Ewen (2018). “Israeli fossils are the oldest modern humans ever found outside of Africa”. Nature 554 (7690): 15–16. Bibcode2018Natur.554...15C. doi:10.1038/d41586-018-01261-5. PMID 29388957. 
  5. ^ a b Zimmer, Carl (2019年7月10日). “A Skull Bone Discovered in Greece May Alter the Story of Human Prehistory”. The New York Times. https://www.nytimes.com/2019/07/10/science/skull-neanderthal-human-europe-greece.html 2019年12月31日閲覧。 
  6. ^ a b Staff (2019年7月10日). “'Oldest remains' outside Africa reset human migration clock”. Phys.org. https://phys.org/news/2019-07-oldest-africa-reset-human-migration.html 2019年7月10日閲覧。 
  7. ^ a b Delson, Eric (10 July 2019). “An early dispersal of modern humans from Africa to Greece – Analysis of two fossils from a Greek cave has shed light on early hominins in Eurasia. One fossil is the earliest known specimen of Homo sapiens found outside Africa; the other is a Neanderthal who lived 40,000 years later.”. Nature 571 (7766): 487–488. doi:10.1038/d41586-019-02075-9. PMID 31337897. 
  8. ^ “Special Feature: The Neandertal Genome” (英語). Science (AAAS). (5 2010). http://www.sciencemag.org/special/neandertal/ 2010年8月12日閲覧。.  アブストラクト和訳 (PDF)
  9. ^ Lohse, K; Frantz, L.A.F. (2014). “Neandertal Admixture in Eurasia Confirmed by Maximum-Likelihood Analysis of Three Genomes”. Genetics 196 (4): 1241–1251. doi:10.1534/genetics.114.162396. PMC 3982695. PMID 24532731. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3982695/. 
  10. ^ Reich, D; Green, RE; Kircher, M; Krause, J; Patterson, N; Durand, EY; Viola, B; Briggs, AW et al. (2010). “Genetic history of an archaic hominin group from Denisova Cave in Siberia”. Nature 468 (7327): 1053–1060. Bibcode2010Natur.468.1053R. doi:10.1038/nature09710. PMC 4306417. PMID 21179161. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4306417/. 
  11. ^ Hammer, M.F.; Woerner, A.E.; Mendez, F.L.; Watkins, J.C.; Wall, J.D. (2011). “Genetic evidence for archaic admixture in Africa”. Proceedings of the National Academy of Sciences 108 (37): 15123–15128. Bibcode2011PNAS..10815123H. doi:10.1073/pnas.1109300108. PMC 3174671. PMID 21896735. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3174671/. 
  12. ^ Richter, David (8 June 2017). “The age of the hominin fossils from Jebel Irhoud, Morocco, and the origins of the Middle Stone Age”. Nature 546 (7657): 293–296. Bibcode2017Natur.546..293R. doi:10.1038/nature22335. PMID 28593967. 
  13. ^ “Earliest evidence of modern human life history in North African early Homo sapiens”. Proceedings of the National Academy of Sciences of the United States of America 104 (15): 6128–33. (April 2007). Bibcode2007PNAS..104.6128S. doi:10.1073/pnas.0700747104. PMC 1828706. PMID 17372199. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1828706/. 
  14. ^ Mcdougall, I.; Brown, H.; Fleagle, G. (Feb 2005). “Stratigraphic placement and age of modern humans from Kibish, Ethiopia”. Nature 433 (7027): 733–736. Bibcode2005Natur.433..733M. doi:10.1038/nature03258. ISSN 0028-0836. PMID 15716951. 
  15. ^ Stringer, C. (2016). “The origin and evolution of Homo sapiens”. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences 371 (1698): 20150237. doi:10.1098/rstb.2015.0237. PMC 4920294. PMID 27298468. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4920294/. 
  16. ^ Sample, Ian (2017年6月7日). “Oldest Homo sapiens bones ever found shake foundations of the human story”. The Guardian. https://www.theguardian.com/science/2017/jun/07/oldest-homo-sapiens-bones-ever-found-shake-foundations-of-the-human-story 2017年6月7日閲覧。 
  17. ^ Hublin, Jean-Jacques; Ben-Ncer, Abdelouahed; Bailey, Shara E.; Freidline, Sarah E.; Neubauer, Simon; Skinner, Matthew M.; Bergmann, Inga; Le Cabec, Adeline et al. (2017). “New fossils from Jebel Irhoud, Morocco and the pan-African origin of Homo sapiens. Nature 546 (7657): 289–292. Bibcode2017Natur.546..289H. doi:10.1038/nature22336. PMID 28593953. http://kar.kent.ac.uk/62267/1/Submission_288356_1_art_file_2637492_j96j1b.pdf. 
  18. ^ Scerri, Eleanor M. L.; Thomas, Mark G.; Manica, Andrea; Gunz, Philipp; Stock, Jay T.; Stringer, Chris; Grove, Matt; Groucutt, Huw S. et al. (2018). “Did Our Species Evolve in Subdivided Populations across Africa, and Why Does It Matter?”. Trends in Ecology & Evolution 33 (8): 582–594. doi:10.1016/j.tree.2018.05.005. PMC 6092560. PMID 30007846. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6092560/. 
  19. ^ Zimmer, Carl (2019年9月10日). “Scientists Find the Skull of Humanity's Ancestor, on a Computer – By comparing fossils and CT scans, researchers say they have reconstructed the skull of the last common forebear of modern humans.”. The New York Times. https://www.nytimes.com/2019/09/10/science/human-ancestor-skull-computer.html 2019年12月31日閲覧。 
  20. ^ Mounier, Aurélien; Lahr, Marta (2019). “Deciphering African late middle Pleistocene hominin diversity and the origin of our species”. Nature Communications 10 (1): 3406. Bibcode2019NatCo..10.3406M. doi:10.1038/s41467-019-11213-w. PMC 6736881. PMID 31506422. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6736881/. 
  21. ^ a b “The earliest modern humans outside Africa”. Science 359 (6374): 456–459. (26 Jan 2018). Bibcode2018Sci...359..456H. doi:10.1126/science.aap8369. PMID 29371468. 
  22. ^ a b c d e Lopez, Saioa; van Dorp, Lucy; Hellenthal, Garrett (2016). “Human Dispersal Out of Africa: A Lasting Debate”. Evolutionary Bioinformatics 11s2 (Suppl 2): 57–68. doi:10.4137/EBO.S33489. ISSN 1176-9343. PMC 4844272. PMID 27127403. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4844272/. 
  23. ^ a b Scerri, Eleanor (2017). “The Stone Age Archaeology of West Africa”. African History. doi:10.1093/acrefore/9780190277734.013.137. ISBN 9780190277734. https://oxfordre.com/africanhistory/view/10.1093/acrefore/9780190277734.001.0001/acrefore-9780190277734-e-137. 
  24. ^ Rito, T; Richards, MB; Fernandes, V; Alshamali, F; Cerny, V; Pereira, L; Soares, P (2013). “The first modern human dispersals across Africa”. PLOS ONE 8 (11): e80031. Bibcode2013PLoSO...880031R. doi:10.1371/journal.pone.0080031. PMC 3827445. PMID 24236171. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3827445/. 
  25. ^ “Patterns of Ancestry, Signatures of Natural Selection, and Genetic Association with Stature in Western African Pygmies”. PLOS Genetics 8 (4): e1002641. (2012). doi:10.1371/journal.pgen.1002641. PMC 3343053. PMID 22570615. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3343053/. 
  26. ^ “Genetic Variation and Recent Positive Selection in Worldwide Human Populations: Evidence from Nearly 1 Million SNPs”. PLOS ONE 4 (11): e7888. (2009). Bibcode2009PLoSO...4.7888L. doi:10.1371/journal.pone.0007888. PMC 2775638. PMID 19924308. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2775638/. 
  27. ^ “The Genetic Structure and History of Africans and African Americans”. Science 324 (5930): 1035–1044. (2009). Bibcode2009Sci...324.1035T. doi:10.1126/science.1172257. PMC 2947357. PMID 19407144. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2947357/.  (Supplementary Data)
  28. ^ “Maternal traces of deep common ancestry and asymmetric gene flow between Pygmy hunter–gatherers and Bantu-speaking farmers”. PNAS 105 (5): 1596–601. (2008). Bibcode2008PNAS..105.1596Q. doi:10.1073/pnas.0711467105. PMC 2234190. PMID 18216239. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2234190/.  fig. 3.
  29. ^ “Inferring the Demographic History of African Farmers and Pygmy Hunter–Gatherers Using a Multilocus Resequencing Data Set”. PLOS Genetics 5 (4): e1000448. (2009). doi:10.1371/journal.pgen.1000448. PMC 2661362. PMID 19360089. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2661362/. 
  30. ^ “Scientists discover oldest known modern human fossil outside of Africa: Analysis of fossil suggests Homo sapiens left Africa at least 50,000 years earlier than previously thought”. ScienceDaily. https://www.sciencedaily.com/releases/2018/01/180125140923.htm 2018年1月27日閲覧。 
  31. ^ Ghosh, Pallab (2018年). “Modern humans left Africa much earlier”. BBC News. https://www.bbc.co.uk/news/science-environment-42817323 2018年1月27日閲覧。 
  32. ^ The Palaeolithic Jebel Qafzeh Cave in Palestine”. ancientneareast.tripod.com. 2017年11月25日閲覧。
  33. ^ “Human footprints dating back 120,000 years found in Saudi Arabia” (英語). phys.org. https://phys.org/news/2020-09-ancient-footprints-saudi-arabia-humans.html 2020年10月9日閲覧。 
  34. ^ Stewart, Mathew; Clark-Wilson, Richard; Breeze, Paul S.; Janulis, Klint; Candy, Ian; Armitage, Simon J.; Ryves, David B.; Louys, Julien et al. (1 September 2020). “Human footprints provide snapshot of last interglacial ecology in the Arabian interior” (英語). Science Advances 6 (38): eaba8940. Bibcode2020SciA....6.8940S. doi:10.1126/sciadv.aba8940. ISSN 2375-2548. PMC 7500939. PMID 32948582. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7500939/. 
  35. ^ Lawler, Andrew (2011). “Did Modern Humans Travel Out of Africa Via Arabia?”. Science 331 (6016): 387. Bibcode2011Sci...331..387L. doi:10.1126/science.331.6016.387. PMID 21273459. 
  36. ^ Trail of 'Stone Breadcrumbs' Reveals the Identity of One of the First Human Groups to Leave Africa”. ScienceDaily (2011年12月1日). 2019年5月8日閲覧。
  37. ^ Bower, Bruce (2011年1月27日). “Hints of earlier human exit from Africa”. Science News. 2019年5月8日閲覧。
  38. ^ Liu, Wu (2010). “Human remains from Zhirendong, South China, and modern human emergence in East Asia”. Proceedings of the National Academy of Sciences 107 (45): 19201–19206. Bibcode2010PNAS..10719201L. doi:10.1073/pnas.1014386107. PMC 2984215. PMID 20974952. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2984215/. 
  39. ^ Dennell, Robin (2010). “Two interpretations of the Zhirendong mandible described by Liu and colleagues”. Nature 468 (7323): 512–513. doi:10.1038/468512a. PMID 21107416. 
  40. ^ Shena, Guanjun (2002). “U-Series dating of Liujiang hominid site in Guangxi, Southern China”. Journal of Human Evolution 43 (6): 817–829. doi:10.1006/jhev.2002.0601. PMID 12473485. 
  41. ^ Lunadong fossils support theory of earlier dispersal of modern man”. University of Hawaii at Mānoa (2014年9月18日). 2019年5月8日閲覧。
  42. ^ a b Ahmed Sayeed (2020-01-15). Negate Fighting Faith. Sankalp Publication. pp. 115 
  43. ^ Harpending, Henry; Cochran, Gregory. The 10,000 Year Explosion. Basic Books. p. 214. ISBN 978-0-465-00221-4. https://lesacreduprintemps19.files.wordpress.com/2012/12/the-10000-year-explosion-how-civilization-accelerated-human-evolution-2009-by-gregory-cochran-henry-harpending.pdf 2015年12月1日閲覧。 
  44. ^ Ambrose, Stanley (1998). “Late Pleistocene human population bottlenecks, volcanic winter, and differentiation of modern humans”. Journal of Human Evolution 34 (6): 623–651. doi:10.1006/jhev.1998.0219. PMID 9650103. 
  45. ^ Balter, Michael (2010). “Of Two Minds About Toba's Impact”. Science 327 (5970): 1187–1188. Bibcode2010Sci...327.1187B. doi:10.1126/science.327.5970.1187-a. PMID 20203021. 
  46. ^ Karmin M, Saag L, Vicente M, Wilson Sayres MA, Järve M, Talas UG, et al. (April 2015). "A recent bottleneck of Y chromosome diversity coincides with a global change in culture". Genome Research. 25 (4): 459–66. doi:10.1101/gr.186684.114. PMC 4381518. PMID 25770088
  47. ^ a b “A Rare Deep-Rooting D0 African Y-chromosomal Haplogroup and its Implications for the Expansion of Modern Humans Out of Africa”. Genetics 212 (4): 1421–1428. (June 2019). doi:10.1534/genetics.119.302368. PMC 6707464. PMID 31196864. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6707464/. 
  48. ^ Zhivotovsky, Lev A; Rosenberg, Noah A; Feldman, Marcus W (2003). “Features of Evolution and Expansion of Modern Humans, Inferred from Genomewide Microsatellite Markers”. American Journal of Human Genetics 72 (5): 1171–1186. doi:10.1086/375120. PMC 1180270. PMID 12690579. https://doi.org/10.1086/375120. 
  49. ^ Stix, Gary (2008年). “The Migration History of Humans: DNA Study Traces Human Origins Across the Continents”. 2011年6月14日閲覧。
  50. ^ Kevin O. Pope; John E. Terrell (9 October 2007), “Environmental setting of human migrations in the circum-Pacific region”, Journal of Biogeography 35 (1): 071009214220006––, doi:10.1111/j.1365-2699.2007.01797.x 
  51. ^ Vincent Macaulay (13 May 2005), “Single, Rapid Coastal Settlement of Asia Revealed by Analysis of Complete Mitochondrial Genomes; Vol. 308. no. 5724”, Science Magazine 308 (5724): 1034–36, doi:10.1126/science.1109792, PMID 15890885, http://sciencemag.org/cgi/content/abstract/308/5724/1034 
  52. ^ “Pleistocene Mitochondrial Genomes Suggest a Single Major Dispersal of Non-Africans and a Late Glacial Population Turnover in Europe”. Current Biology 26 (6): 827–833. (2016). doi:10.1016/j.cub.2016.01.037. hdl:2440/114930. PMID 26853362. 
  53. ^ “Ancestral mitochondrial N lineage from the Neolithic 'green' Sahara”. Scientific Reports 9 (1): 3530. (March 2019). Bibcode2019NatSR...9.3530V. doi:10.1038/s41598-019-39802-1. PMC 6401177. PMID 30837540. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6401177/. 
  54. ^ Metspalu, M; Kivisild, T; Metspalu, E; Parik, J; Hudjashov, G; Kaldma, K; Serk, P; Karmin, M; Behar, DM; Gilbert, M Thomas P; Endicott, Phillip; Mastana, Sarabjit; Papiha, Surinder S; Skorecki, Karl; Torroni, Antonio; Villems, Richard; et al. (2004). "Most of the extant mtDNA boundaries in South and Southwest Asia were likely shaped during the initial settlement of Eurasia by anatomically modern humans". BMC Genetics. 5: 26. doi:10.1186/1471-2156-5-26. PMC 516768. PMID 15339343
  55. ^ Fregel, R; Cabrera, V; Larruga, JM; Abu-Amero, KK; González, AM (2015). “Carriers of Mitochondrial DNA Macrohaplogroup N Lineages Reached Australia around 50,000 Years Ago following a Northern Asian Route”. PLOS ONE 10 (6): e0129839. Bibcode2015PLoSO..1029839F. doi:10.1371/journal.pone.0129839. PMC 4460043. PMID 26053380. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4460043/. 
  56. ^ Dennell, Robin; Petraglia, Michael D. (2012). “The dispersal of Homo sapiens across southern Asia: how early, how often, how complex?”. Quaternary Science Reviews 47: 15–22. Bibcode2012QSRv...47...15D. doi:10.1016/j.quascirev.2012.05.002. 
  57. ^ Prüfer, Kay (2013). “The complete genome sequence of a Neanderthal from the Altai Mountains”. Nature 505 (7481): 43–49. Bibcode2014Natur.505...43P. doi:10.1038/nature12886. PMC 4031459. PMID 24352235. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4031459/. 
  58. ^ a b c Bowler, James M. (2003). “New ages for human occupation and climatic change at Lake Mungo, Australia”. Nature 421 (6925): 837–840. Bibcode2003Natur.421..837B. doi:10.1038/nature01383. PMID 12594511. 
  59. ^ Wood, Rachel (2017-09-02). “Comments on the chronology of Madjedbebe”. Australian Archaeology 83 (3): 172–174. doi:10.1080/03122417.2017.1408545. ISSN 0312-2417. 
  60. ^ “Homo sapiens first reach Southeast Asia and Sahul?”. Proceedings of the National Academy of Sciences of the United States of America 115 (34): 8482–8490. (August 2018). doi:10.1073/pnas.1808385115. PMC 6112744. PMID 30082377. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6112744/. 
  61. ^ First Mariners – Archaeology Magazine Archive. Archive.archaeology.org. Retrieved on 2013-11-16.
  62. ^ Callaway, Ewen (22 September 2011), “First Aboriginal genome sequenced”, Nature, doi:10.1038/news.2011.551, http://www.nature.com/news/2011/110922/full/news.2011.551.html 
  63. ^ Reich (2011), “Denisova Admixture and the First Modern Human Dispersals into Southeast Asia and Oceania”, The American Journal of Human Genetics 89 (4): 516–528, doi:10.1016/j.ajhg.2011.09.005, PMC 3188841, PMID 21944045, http://www.pubmedcentral.nih.gov/articlerender.fcgi?tool=pmcentrez&artid=3188841 
  64. ^ Choi, Charles (2011年9月22日). “Now-Extinct Relative Had Sex with Humans Far and Wide”. LiveScience. 2021年9月13日閲覧。
  65. ^ Pleistocene Sea Level Maps”. Fieldmuseum.org. 2010年9月23日閲覧。
  66. ^ Rasmussen, M (Oct 2011). “An Aboriginal Australian genome reveals separate human dispersals into Asia”. Science 334 (6052): 94–98. Bibcode2011Sci...334...94R. doi:10.1126/science.1211177. PMC 3991479. PMID 21940856. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3991479/. 
  67. ^ “Revealing the prehistoric settlement of Australia by Y chromosome and mtDNA analysis”. Proc Natl Acad Sci USA 104 (21): 8726–8730. (May 2007). Bibcode2007PNAS..104.8726H. doi:10.1073/pnas.0702928104. PMC 1885570. PMID 17496137. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1885570/. 
  68. ^ Wade, Nicholas (2007年5月8日). “From DNA Analysis, Clues to a Single Australian Migration”. Nytimes.com (Australia). https://www.nytimes.com/2007/05/08/science/08abor.html 2019年12月31日閲覧。 
  69. ^ Clarkson, Chris; Smith, Mike; Marwick, Ben; Fullagar, Richard; Wallis, Lynley A.; Faulkner, Patrick; Manne, Tiina; Hayes, Elspeth et al. (June 2015). “The archaeology, chronology and stratigraphy of Madjedbebe (Malakunanja II): A site in northern Australia with early occupation”. Journal of Human Evolution 83: 46–64. doi:10.1016/j.jhevol.2015.03.014. hdl:1773/33254. PMID 25957653. https://ro.uow.edu.au/smhpapers/3027. 
  70. ^ Clarkson, Chris; Jacobs, Zenobia; Marwick, Ben; Fullagar, Richard; Wallis, Lynley; Smith, Mike; Roberts, Richard G.; Hayes, Elspeth et al. (19 July 2017). “Human occupation of northern Australia by 65,000 years ago”. Nature 547 (7663): 306–310. Bibcode2017Natur.547..306C. doi:10.1038/nature22968. hdl:2440/107043. PMID 28726833. https://hdl.handle.net/2440/107043. 
  71. ^ Flannery, Tim (2002), "The Future Eaters: An Ecological History of the Australasian Lands and People" (Grove Press)
  72. ^ “Fossil Teeth Put Humans in Europe Earlier Than Thought”. The New York Times. (2011年11月2日). https://www.nytimes.com/2011/11/03/science/fossil-teeth-put-humans-in-europe-earlier-than-thought.html 2019年12月31日閲覧。 
  73. ^ Oppenheimer, Stephen "Out of Eden: Peopling of the World" (Robinson; New Ed edition (1 March 2012))
  74. ^ Benazzi, S.; Douka, K.; Fornai, C.; Bauer, C.C.; Kullmer, O.; Svoboda, J.F.; Pap, I.; Mallegni, F. et al. (2011). “Early dispersal of modern humans in Europe and implications for Neanderthal behaviour”. Nature 479 (7374): 525–528. Bibcode2011Natur.479..525B. doi:10.1038/nature10617. PMID 22048311. 
  75. ^ Higham, T.; Compton, T.; Stringer, C.; Jacobi, R.; Shapiro, B.; Trinkaus, E.; Chandler, B.; Gröning, F. et al. (2011). “The earliest evidence for anatomically modern humans in northwestern Europe”. Nature 479 (7374): 521–524. Bibcode2011Natur.479..521H. doi:10.1038/nature10484. PMID 22048314. 
  76. ^ Sankararaman, S.; Patterson, N.; Li, H.; Pääbo, S.; Reich, D; Akey, J.M. (2012). “The Date of Interbreeding between Neandertals and Modern Humans”. PLOS Genetics 8 (10): e1002947. arXiv:1208.2238. Bibcode2012arXiv1208.2238S. doi:10.1371/journal.pgen.1002947. PMC 3464203. PMID 23055938. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3464203/. 
  77. ^ a b Hoffecker, J. (2006). A Prehistory of the North: Human Settlements of the Higher Latitudes. New Jersey: Rutgers University Press. p. 101. https://archive.org/details/prehistorynorthh00hoff 
  78. ^ Hoffecker, John F. (2002). Desolate landscapes: Ice-Age settlement in Eastern Europe. New Brunswick: Rutgers University Press. pp. 158–162, 217–233 
  79. ^ a b Pavlov, P; Svendsen, JI; Indrelid, S (2001). “Human presence in the European Arctic nearly 40,000 years ago”. Nature 413 (6851): 64–67. Bibcode2001Natur.413...64P. doi:10.1038/35092552. PMID 11544525. 
  80. ^ Mamontovaya Kurya:an enigmatic, nearly 40000 years old Paleolithic site in the Russian Arctic”. 2021年9月13日閲覧。
  81. ^ Atlas of human journey: 45–40,000”. The genographic project. National Geographic Society (1996–2010). 2019年5月8日閲覧。
  82. ^ Caramelli, D; Lalueza-Fox, C; Vernesi, C; Lari, M; Casoli, A; Mallegni, F; Chiarelli, B; Dupanloup, I et al. (May 2003). “Evidence for a genetic discontinuity between Neandertals and 24,000-year-old anatomically modern Europeans”. PNAS 100 (11): 6593–6597. Bibcode2003PNAS..100.6593C. doi:10.1073/pnas.1130343100. ISSN 0027-8424. PMC 164492. PMID 12743370. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC164492/. 
  83. ^ Currat, M.; Excoffier, L. (2004). "Modern Humans Did Not Admix with Neanderthals during Their Range Expansion into Europe". PLoS Biol. 2 (12): e421. PMC 532389 Freely accessible. PMID 15562317. doi:10.1371/journal.pbio.0020421.
  84. ^ “Major genomic mitochondrial lineages delineate early human expansions”. BMC Genet. 2 (1): 13. (2001). doi:10.1186/1471-2156-2-13. PMC 55343. PMID 11553319. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC55343/. 
  85. ^ “Modern humans did not admix with Neanderthals during their range expansion into Europe”. PLOS Biology 2 (12): e421. (Dec 2004). doi:10.1371/journal.pbio.0020421. PMC 532389. PMID 15562317. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC532389/. 
  86. ^ Neanderthals disappeared from Europe earlier than thought, says study”. phys.org (2021年3月8日). 2021年9月13日閲覧。
  87. ^ Neanderthals Died Out 10,000 Years Earlier Than Thought, With Help From Modern Humans”. nationalgeographic (2014年8月21日). 2021年9月13日閲覧。
  88. ^ Yang (2017). “40,000-Year-Old Individual from Asia Provides Insight into Early Population Structure in Eurasia”. Current Biology 27 (20): 3202–3208. doi:10.1016/j.cub.2017.09.030. PMC 6592271. PMID 29033327. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6592271/. 
  89. ^ Ding, Q.; Hu, Y.; Xu, S.; Wang, J.; Jin, L. (2014). “Neanderthal Introgression at Chromosome 3p21.31 was Under Positive Natural Selection in East Asians”. Molecular Biology and Evolution 31 (3): 683–695. doi:10.1093/molbev/mst260. PMID 24336922. 
  90. ^ Downloadable genotypes of present-day and ancient DNA data (compiled from published papers) | David Reich Lab”. reich.hms.harvard.edu. 2019年9月11日閲覧。
  91. ^ Jeong (Jan 2016). “Deep History of East Asian Populations Revealed Through Genetic Analysis of the Ainu”. Genetics 202 (1): 261–272. doi:10.1534/genetics.115.178673. PMC 4701090. PMID 26500257. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4701090/. 
  92. ^ Kamberov, Yana G (14 February 2013). “Modeling Recent Human Evolution in Mice by Expression of a Selected EDAR Variant”. Cell 152 (4): 691–702. doi:10.1016/j.cell.2013.01.016. PMC 3575602. PMID 23415220. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3575602/. 
  93. ^ Wade, Nicholas (2013年2月14日). “East Asian Physical Traits Linked to 35,000-Year-Old Mutation”. The New York Times. https://www.nytimes.com/2013/02/15/science/studying-recent-human-evolution-at-the-genetic-level.html 2019年12月31日閲覧。 
  94. ^ Peng, Y (2010). “The ADH1B ARG47His polymorphism in East Asian populations and expansion of rice domestication in history”. BMC Evolutionary Biology 10 (15): 15. doi:10.1186/1471-2148-10-15. PMC 2823730. PMID 20089146. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2823730/. 
  95. ^ a b Refugia”. Encyclopedia of Earth. 2012年7月11日閲覧。
  96. ^ Pitulko, V.V.; Nikolsky, P.A.; Girya, E.Y.; Basilyan, A.E.; Tumskoy, V.E.; Koulakov, S.A.; Astakhov, S.N.; Pavlova, E.Y. et al. (2004). “The Yana RHS Site: Humans in the Arctic Before the Last Glacial Maximum”. Science 303 (5654): 52–56. Bibcode2004Sci...303...52P. doi:10.1126/science.1085219. PMID 14704419. 
  97. ^ a b Wells, Spencer; Read, Mark (2002). The Journey of Man – A Genetic Odyssey. Random House. pp. 138–140. ISBN 978-0-8129-7146-0. https://books.google.com/books?id=WAsKm-_zu5sC&pg=PA138 
  98. ^ a b Paleoamerican”. Smithsonian Institution Anthropology Outreach Office. 2009年1月5日時点のオリジナルよりアーカイブ。2009年1月15日閲覧。
  99. ^ A DNA Search for the First Americans Links Amazon Groups to Indigenous Australians”. Smithsonian Institution. 2019年5月8日閲覧。
  100. ^ Bonatto, S. L.; Salzano, F. M. (1997). “A single and early migration for the peopling of greater America supported by mitochondrial DNA sequence data”. Proceedings of the National Academy of Sciences 94 (5): 1866–1871. Bibcode1997PNAS...94.1866B. doi:10.1073/pnas.94.5.1866. PMC 20009. PMID 9050871. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC20009/. 
  101. ^ a b Atlas of the Human Journey-The Genographic Project”. National Geographic Society (1996–2008). 2011年5月1日時点のオリジナルよりアーカイブ。2017年1月27日閲覧。
  102. ^ The peopling of the Americas: Genetic ancestry influences health”. Scientific American. 2019年5月8日閲覧。
  103. ^ Fladmark, K. R. (1979). “Alternate Migration Corridors for Early Man in North America”. American Antiquity 44 (1): 55–69. doi:10.2307/279189. JSTOR 279189. 
  104. ^ 68 Responses to "Sea will rise 'to levels of last Ice Age'"”. Center for Climate Systems Research, Columbia University. 2009年11月17日閲覧。
  105. ^ Su, Bing; Ma, Runlin Z.; Zhang, Xiaoming; Peng, Yi; Zhong, Hua; Qi, Xuebin; Shi, Hong (20 June 2013). “Genetic Evidence of an East Asian Origin and Paleolithic Northward Migration of Y-chromosome Haplogroup N” (英語). PLOS ONE 8 (6): e66102. Bibcode2013PLoSO...866102S. doi:10.1371/journal.pone.0066102. ISSN 1932-6203. PMC 3688714. PMID 23840409. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3688714/. 
  106. ^ Gibbons, Ann (2014年9月4日). “Three-part ancestry for Europeans”. Science. American Association for the Advancement of Science. 2014年10月11日時点のオリジナルよりアーカイブ。2014年10月30日閲覧。
  107. ^ Busby, George BJ; Band, Gavin; Si Le, Quang; Jallow, Muminatou; Bougama, Edith; Mangano, Valentina D; Amenga-Etego, Lucas N; Enimil, Anthony et al. (2016). “Admixture into and within sub-Saharan Africa”. eLife 5. doi:10.7554/eLife.15266. ISSN 2050-084X. PMC 4915815. PMID 27324836. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4915815/. 
  108. ^ Ramsay, Michèle; Hazelhurst, Scott; Sengupta, Dhriti; Aron, Shaun; Choudhury, Ananyo (2018-08-01). “African genetic diversity provides novel insights into evolutionary history and local adaptations” (英語). Human Molecular Genetics 27 (R2): R209–R218. doi:10.1093/hmg/ddy161. ISSN 0964-6906. PMC 6061870. PMID 29741686. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6061870/. 
  109. ^ a b Meacham, William (1984–1985). “On the improbability of Austronesian origins in South China”. Asian Perspective 26: 89–106. JSTOR 42928108. https://www.jstor.org/stable/42928108. 
  110. ^ a b Bellwood, Peter (1991). “The Austronesian Dispersal and the Origin of Languages”. Scientific American 265 (1): 88–93. Bibcode1991SciAm.265a..88B. doi:10.1038/scientificamerican0791-88. JSTOR 24936983. 
  111. ^ Heath, Helen; Summerhayes, Glenn R.; Hung, Hsiao-chun (2017). “Enter the Ceramic Matrix: Identifying the Nature of the Early Austronesian Settlement in the Cagayan Valley, Philippines”. In Piper, Philip J.; Matsumara, Hirofumi; Bulbeck, David. New Perspectives in Southeast Asian and Pacific Prehistory. terra australis. 45. ANU Press. ISBN 9781760460952. http://press-files.anu.edu.au/downloads/press/n2320/html/ch12.xhtml?referer=&page=19 
  112. ^ Carson, Mike T.; Hung, Hsiao-chun; Summerhayes, Glenn; Bellwood, Peter (January 2013). “The Pottery Trail From Southeast Asia to Remote Oceania”. The Journal of Island and Coastal Archaeology 8 (1): 17–36. doi:10.1080/15564894.2012.726941. hdl:1885/72437. 
  113. ^ Madagascar Founded By Women”. Discovery.com. 2012年3月23日閲覧。
  114. ^ Murray P. Cox; Michael G. Nelson; Meryanne K. Tumonggor; François-X. Ricaut; Herawati Sudoyo (March 21, 2012). “A small cohort of Island Southeast Asian women founded Madagascar”. Proceedings of the Royal Society B 279 (1739): 2761–2768. doi:10.1098/rspb.2012.0012. PMC 3367776. PMID 22438500. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3367776/. 
  115. ^ Wanita Indonesia Nenek Moyang Penduduk Madagaskar”. Yahoo News Indonesia (2012年3月21日). 2012年3月23日閲覧。
  116. ^ Did a butterfly effect change the history of the Pacific?”. Stockholm Resilience Centre. Stockholm University. 2019年11月9日閲覧。
  117. ^ Boomert, Arie, 1946- (2016). The indigenous peoples of Trinidad and Tobago : from the first settlers until today. Leiden. pp. 15. ISBN 978-90-8890-354-0. OCLC 944910446 
  118. ^ Nägele, Kathrin; Posth, Cosimo; Iraeta Orbegozo, Miren; Chinique de Armas, Yadira; Hernández Godoy, Silvia Teresita; González Herrera, Ulises M.; Nieves-Colón, Maria A.; Sandoval-Velasco, Marcela et al. (2020). “Genomic insights into the early peopling of the Caribbean” (英語). Science 369 (6502): 456–460. doi:10.1126/science.aba8697. ISSN 0036-8075. PMID 32499399. 
  119. ^ "Prehistory of the Caribbean Culture Area", Southeast Archaeological Center, National Park Service (retrieved 19 July 2013)
  120. ^ Keegan, William F., 1955- (2017). The Caribbean before Columbus. Hofman, Corinne L., 1959-. New York, NY. pp. 51. ISBN 978-0-19-060524-7. OCLC 949669477 
  121. ^ Fitzpatrick, Scott M. (2013-03-07). “The Southward Route Hypothesis”. Oxford Handbooks Online: 202. doi:10.1093/oxfordhb/9780195392302.013.0068. 
  122. ^ Fagan, B.M. (2007). People of the earth: An introduction to world prehistory. Upper Saddle River, NJ: Pearson Prentice Hall
  123. ^ Napolitano, Matthew F.; DiNapoli, Robert J.; Stone, Jessica H.; Levin, Maureece J.; Jew, Nicholas P.; Lane, Brian G.; O’Connor, John T.; Fitzpatrick, Scott M. (2019). “Reevaluating human colonization of the Caribbean using chronometric hygiene and Bayesian modeling”. Science Advances 5 (12): 7. doi:10.1126/sciadv.aar7806. ISSN 2375-2548. PMC 6957329. PMID 31976370. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6957329/. 
  124. ^ Hoffecker, John F. (2005). A prehistory of the north: human settlement of the higher latitudes. Rutgers University Press. p. 130. ISBN 978-0-8135-3469-5. https://books.google.com/books?id=_rL5F4EAaFkC&pg=PA132 
  125. ^ Gibbon, pp. 28–31
  126. ^ Rigby, Bruce. “101. Qaummaarviit Historic Park, Nunavut Handbook”. 2006年5月29日時点のオリジナルよりアーカイブ。2009年10月2日閲覧。
  127. ^ Wood, Shannon Raye (1992-04). Tooth wear and the sexual division of labour in an Inuit population. Simon Fraser University. https://summit.sfu.ca/item/3743 2024年1月29日閲覧。. 
  128. ^ “New Study Offers Clues to Swift Arctic Extinction”. The New York Times. (2014年8月29日). https://www.nytimes.com/2014/08/29/science/study-offers-clues-to-arctic-mystery-paleo-eskimos-abrupt-extinction.html 
  129. ^ You, Jia (2014年8月28日). “The strange history of the North American Arctic” (英語). Science. https://www.sciencemag.org/news/2014/08/strange-history-north-american-arctic 





英和和英テキスト翻訳>> Weblio翻訳
英語⇒日本語日本語⇒英語
  
  •  現生人類の拡散のページへのリンク

辞書ショートカット

すべての辞書の索引

「現生人類の拡散」の関連用語

現生人類の拡散のお隣キーワード
検索ランキング

   

英語⇒日本語
日本語⇒英語
   



現生人類の拡散のページの著作権
Weblio 辞書 情報提供元は 参加元一覧 にて確認できます。

   
ウィキペディアウィキペディア
All text is available under the terms of the GNU Free Documentation License.
この記事は、ウィキペディアの現生人類の拡散 (改訂履歴)の記事を複製、再配布したものにあたり、GNU Free Documentation Licenseというライセンスの下で提供されています。 Weblio辞書に掲載されているウィキペディアの記事も、全てGNU Free Documentation Licenseの元に提供されております。

©2024 GRAS Group, Inc.RSS